[1] Brocks J J, Okenane Schaeffer P., a biomarker for purple sulfur bacteria (Chromatiaceae), and other new carotenoid derivatives from the1640Ma Barney Creek Formation[J]. Geochimica et Cosmochimica Acta, 2008, 72(5): 1396-1414.
[2] 马健,吴朝东,王熠哲,等. 准噶尔盆地渐新世安集海河组类胡萝卜素的发现及古环境意义[J]. 地质学报,2020,94(6):1853-1868.

Ma Jian, Wu Chaodong, Wang Yizhe, et al. Discovery of carotenoids and its paleolake significance in the Oligocene Anjihaihe Formation, southern Junggar Basin, China[J]. Acta Geologica Sinica, 2020, 94(6): 1853-1868.
[3] Anders D E, Robinson W E. Cycloalkane constituents of the bitumen from Green River Shale[J]. Geochimica et Cosmochimica Acta, 1971, 35(7): 661-678.
[4] Murphy M T J, McCormick A, Eglinton G. Perhydro-β-carotene in the Green River Shale[J]. Science, 1967, 157(3792): 1040-1042.
[5] Lee C, Brocks J J. Identification of carotane breakdown products in the 1.64 billion year old Barney Creek Formation, McArthur Basin, northern Australia[J]. Organic Geochemistry, 2011, 42(4): 425-430.
[6] Casilli A, Silva R C, Laakia J, et al. High resolution molecular organic geochemistry assessment of Brazilian lacustrine crude oils[J]. Organic Geochemistry, 2014, 68: 61-70.
[7] 王国彬,王熠,李二庭,等. 准噶尔盆地玛湖凹陷西斜坡百口泉组含油储集岩分子与碳同位素地球化学特征及其意义[J]. 地球化学,2017,46(3):276-291.

Wang Guobin, Wang Yi, Li Erting, et al. Molecular and carbon isotopic compositions of oil components in the Baikouquan Formation oil-bearing reservoir rocks on the western slope of the Mahu Sag, Junggar Basin[J]. Geochimica, 2017, 46(3): 276-291.
[8] 蒋文龙,阿布力米提•依明,卞保力,等. 准噶尔盆地西北缘风城组烃源岩热演化生物标志化合物变化及意义[J]. 新疆石油地质,2022,43(6):684-692.

Jiang Wenlong, Ablimit·Yiming, Bian Baoli, et al. Changes and significance of biomarkers in thermal evolution of Fengcheng Formation source rocks in northwestern margin of Junggar Basin[J]. Xinjiang Petroleum Geology, 2022, 43(6): 684-692.
[9] 曾文人,陈璇,付国斌,等. 准噶尔盆地石树沟凹陷二叠系平地泉组烃源岩地球化学特征及形成条件[J]. 东北石油大学学报,2023,47(2):70-80.

Zeng Wenren, Chen Xuan, Fu Guobin, et al. Geochemical characteristics and formation conditions of Permian Pingdiquan Formation source rocks in Shishugou Sag, Junggar Basin[J]. Journal of Northeast Petroleum University, 2023, 47(2): 70-80.
[10] Zhang Z D, Gu Y L, Jin J, et al. Assessing source and maturity of oils in the Mahu Sag, Junggar Basin: Molecular concentrations, compositions and carbon isotopes[J]. Marine and Petroleum Geology, 2022, 141: 105724.
[11] Liu Y Z, Zeng J H, Jin J, et al. Geochemical evaluation of produced petroleum from the Middle Permian Lucaogou reservoirs Junggar Basin and its implication for the unconventional shale oil play[J]. Journal of Petroleum Science and Engineering, 2022, 211: 110202.
[12] Li W W, Cao J, Hu W X, et al. Revealing continuous hydrocarbon generation and shale oil accumulation of saline lacustrine source rocks by sequential extraction[J]. Fuel, 2023, 352: 129081.
[13] Xia L W, Cao J, Bian L Z, et al. Co-evolution of paleo-environment and bio-precursors in a Permian alkaline lake, Mahu mega-oil province, Junggar Basin: Implications for oil sources[J]. Science China Earth Sciences, 2022, 65(3): 462-476.
[14] Ma J, French K L, Cui X Q, et al. Carotenoid biomarkers in Namibian shelf sediments: Anoxygenic photosynthesis during sulfide eruptions in the Benguela upwelling system[J]. Proceedings of the National Academy of Sciences of the United States of America, 2021, 118(29): e2106040118.
[15] Moldowan J M, Seifert W K, Gallegos E J. Relationship between petroleum composition and depositional environment of petroleum source rocks[J]. AAPG Bulletin, 1985, 69(8): 1255-1268.
[16] Ding X J, Gao C H, Zha M, et al. Depositional environment and factors controlling β-carotane accumulation: A case study from the Jimsar Sag, Junggar Basin, northwestern China[J]. Palaeogeography, Palaeoclimatology, Palaeoecology, 2017, 485: 833-842.
[17] Zhang T, Wang Z D, Wang X F, et al. Characterization of oxygen-bearing geolipids in the Upper Permian Lucaogou shale, Santanghu Basin, NE China, using stepwise pyrolysis and hydrous pyrolysis[J]. Marine and Petroleum Geology, 2021, 126: 104926.
[18] 张婷,王作栋,钱宇,等. 长直链烷基苯及其异构体母质来源研究及地球化学意义[J]. 沉积学报,2017,35(4):843-852.

Zhang Ting, Wang Zuodong, Qian Yu, et al. Distribution characteristics of long-chain alkylbenzene and its isomers[J]. Acta Sedimentologica Sinica, 2017, 35(4): 843-852.
[19] Wang Z D, Liang M L, Qian Y, et al. The oxygen-bearing geolipids in the Lucaogou shale of Upper Permian, Santanghu Basin, China[J]. Organic Geochemistry, 2016, 102: 59-66.
[20] 肖洪,李美俊,王铁冠,等. 中元古界沉积物中典型分子标志化合物及其地质意义:以宣隆坳陷下马岭组黑色页岩为例[J]. 沉积学报,2022,40(2):547-556.

Xiao Hong, Li Meijun, Wang Tieguan, et al. Typical molecular marker assemblage of the Mesoproterozoic sediments: A case study of the Xiamaling Formation black shales in the Xuanlong Depression[J]. Acta Sedimentologica Sinica, 2022, 40(2): 547-556.
[21] Xie Z B, Tao H F, Qu Y Q, et al. Synergistic evolution of palaeoenvironment-bionts and hydrocarbon generation of Permian saline lacustrine source rocks in Jimusar Sag, Junggar Basin[J]. Energies, 2023, 16(9): 3797.
[22] Peters K E, Walters C C, Moldowan J M. The biomarker guide: Volume 2: Biomarkers and isotopes in petroleum systems and earth history[M]. New York: Cambridge University Press, 2005.
[23] 徐银波,李锋,张家强,等. 三塘湖盆地石头梅地区二叠系芦草沟组有机质富集特征[J]. 地质学报,2022,96(11):4010-4022.

Xu Yinbo, Li Feng, Zhang Jiaqiang, et al. Enrichment characteristics of organic matter in the Permian Lucaogou Formation in Shitoumei area, Santanghu Basin[J]. Acta Geologica Sinica, 2022, 96(11): 4010-4022.
[24] 赵泽辉,郭召杰,张臣,等. 新疆东部三塘湖盆地构造演化及其石油地质意义[J]. 北京大学学报(自然科学版),2003,39(2):219-228.

Zhao Zehui, Guo Zhaojie, Zhang Chen, et al. Tectonic evolution of the Santanghu Basin, east Xinjiang and its implication for the hydrocarbon accumulation[J]. Acta Scientiarum Naturalium Universitatis Pekinensis, 2003, 39(2): 219-228.
[25] 王作栋,陶明信,梁明亮,等. 三塘湖盆地上二叠统芦草沟组烃源岩地球化学特征[J]. 沉积学报,2012,30(5):975-982.

Wang Zuodong, Tao Mingxin, Liang Mingliang, et al. Characteristics of organic geochemistry of Lucaogou Formation source rocks, Upper Permian, Santanghu Basin[J]. Acta Sedimentologica Sinica, 2012, 30(5): 975-982.
[26] 李玮,柳益群,董云鹏,等. 新疆三塘湖地区石炭纪火山岩年代学、地球化学及其大地构造意义[J]. 中国科学(D辑):地球科学,2012,42(11):1716-1731.

Li Wei, Liu Yiqun, Dong Yunpeng, et al. The geochemical characteristics, geochronology and tectonic significance of the Carboniferous volcanic rocks of the Santanghu area in northeastern Xinjiang, China[J]. Science China (Seri. D): Earth Sciences, 2012, 42(11): 1716-1731.
[27] 焦悦,吴朝东,王家林,等. 天山东段地区二叠系芦草沟组沉积特征与古环境对比[J]. 古地理学报,2023,25(2):277-293.

Jiao Yue, Wu Chaodong, Wang Jialin, et al. Comparative study on sedimentary characteristics and palaeoenvironment of the Permian Lucaogou Formation in eastern Tianshan Mountains[J]. Journal of Palaeogeography, 2023, 25(2): 277-293.
[28] 柳益群,周鼎武,冯乔,等. 新疆北部二叠系特征及其与三叠系界线划分[J]. 西北大学学报(自然科学版),2006,36(4):615-622.

Liu Yiqun, Zhou Dingwu, Feng Qiao, et al. Characterristics of Permian and the boundary between Permian and Triassic system in the northern Xinjiang[J]. Journal of Northwest University (Natural Science Edition), 2006, 36(4): 615-622.
[29] 方向,郝翠果,刘俊田,等. 新疆三塘湖盆地芦草沟组混积岩型致密油形成条件与富集因素[J]. 地质通报,2023,42(2/3):397-410.

Fang Xiang, Hao Cuiguo, Liu Juntian, et al. Geological conditions and enrichment factors of the Lucaogou Formation diamictite tight oil in Santanghu Basin, Xinjiang[J]. Geological Bulletin of China, 2023, 42(2/3): 397-410.
[30] 潘永帅,黄志龙,郭小波,等. 火山灰影响下的湖相富有机质页岩油成藏条件分析:以三塘湖盆地条湖—马朗凹陷芦草沟组为例[J]. 地质学报,2022,96(3):1053-1068.

Pan Yongshuai, Huang Zhilong, Guo Xiaobo, et al. Analysis of accumulation conditions of lacustrine organic-rich shale oil affected by volcanic ash: A case study of the Lucaogou Formation in the Tiaohu-Malang Sag, Santanghu Basin[J]. Acta Geologica Sinica, 2022, 96(3): 1053-1068.
[31] 范谭广,徐雄飞,范亮,等. 三塘湖盆地二叠系芦草沟组页岩油地质特征与勘探前景[J]. 中国石油勘探,2021,26(4):125-136.

Fan Tanguang, Xu Xiongfei, Fan Liang, et al. Geological characteristics and exploration prospect of shale oil in Permian Lucaogou Formation, Santanghu Basin[J]. China Petroleum Exploration, 2021, 26(4): 125-136.
[32] 梁世君,罗劝生,王瑞,等. 三塘湖盆地二叠系非常规石油地质特征与勘探实践[J]. 中国石油勘探,2019,24(5):624-635.

Liang Shijun, Luo Quansheng, Wang Rui, et al. Geological characteristics and exploration practice of unconventional Permian oil resources in the Santanghu Basin[J]. China Petroleum Exploration, 2019, 24(5): 624-635.
[33] 蒋德鑫,姜正龙,张贺,等. 烃源岩总有机碳含量测井预测模型探讨:以陆丰凹陷文昌组为例[J]. 岩性油气藏,2019,31(6):109-117.

Jiang Dexin, Jiang Zhenglong, Zhang He, et al. Well logging prediction models of TOC content in source rocks: A case of Wenchang Formation in Lufeng Sag[J]. Lithologic Reservoirs, 2019, 31(6): 109-117.
[34] Ding W J, Hou D J, Jiang L, et al. High abundance of carotanes in the brackish-saline lacustrine sediments: A possible cyanobacteria source?[J]. International Journal of Coal Geology, 2020, 219: 103373.
[35] 李志明,徐二社,秦建中,等. 烃源岩评价中的若干问题[J]. 西安石油大学学报(自然科学版),2010,25(6):8-12,27.

Li Zhiming, Xu Ershe, Qin Jianzhong, et al. Some problems on the evaluation of source rock[J]. Journal of Xi'an Shiyou University (Natural Science Edition), 2010, 25(6): 8-12, 27.
[36] 卢双舫,马延伶,曹瑞成,等. 优质烃源岩评价标准及其应用:以海拉尔盆地乌尔逊凹陷为例[J]. 地球科学:中国地质大学学报,2012,37(3):535-544.

Lu Shuangfang, Ma Yanling, Cao Ruicheng, et al. Evaluation criteria of high-quality source rocks and its applications: Taking the Wuerxun Sag in Hailaer Basin as an example[J]. Earth Science: Journal of China University of Geosciences, 2012, 37(3): 535-544.
[37] 赵佳玉,王淑贤, Darin A,等. 新疆阿尔泰全新世双湖沉积物正构烷烃分布及其环境意义[J]. 第四纪研究,2021,41(4):965-975.

Zhao Jiayu, Wang Shuxian, Darin A, et al. N-Alkane distribution and their paleoenvironmental implications during Holocene in lacustrine sediments in Lake Shuang, Xinjiang[J]. Quaternary Sciences, 2021, 41(4): 965-975.
[38] Peters K E, Fraser T H, Amris W, et al. Geochemistry of crude oils from eastern Indonesia[J]. AAPG Bulletin, 1999, 83(12): 1927-1942.
[39] Volkman J K. A review of sterol markers for marine and terrigenous organic matter[J]. Organic Geochemistry, 1986, 9(2): 83-99.
[40] Volkman J K. Sterols in microorganisms[J]. Applied Microbiology and Biotechnology, 2003, 60(5): 495-506.
[41] 王作栋,孟仟祥,陶明信,等. 烃源岩中C19~C29甾烷系列和25-降藿烷系列的检出及其地质意义[J]. 沉积学报,2009,27(1):180-185.

Wang Zuodong, Meng Qianxiang, Tao Mingxin, et al. Identification of C19-C29 steranes and 25-norhopanes in source rock and geological significance[J]. Acta Sedimentologica Sinica, 2009, 27(1): 180-185.
[42] Knoll A H, Summons R E, Waldbauer J R, et al. The geological succession of primary producers in the oceans[M]//Falkowski P G, Knoll A H. Evolution of primary producers in the sea. Boston: Academic Press, 2007: 133-163.
[43] Volkman J K. Sterols and other triterpenoids: Source specificity and evolution of biosynthetic pathways[J]. Organic Geochemistry, 2005, 36(2): 139-159.
[44] Zeng W R, Zhang Z H, Wang B R, et al. Formation mechanism of organic-rich mixed sedimentary rocks in saline lacustrine basin, Permian Lucaogou Formation, Jimsar Sag, Junggar Basin, northwest China[J]. Marine and Petroleum Geology, 2023, 156: 106452.
[45] Qiao J Q, Baniasad A, Zieger L, et al. Paleo-depositional environment, origin and characteristics of organic matter of the Triassic Chang 7 member of the Yanchang Formation throughout the mid-western part of the Ordos Basin, China[J]. International Journal of Coal Geology, 2021, 237: 103636.
[46] Luo G M, Hallmann C, Xie S C, et al. Comparative microbial diversity and redox environments of black shale and stromatolite facies in the Mesoproterozoic Xiamaling Formation[J]. Geochimica et Cosmochimica Acta, 2015, 151: 150-167.
[47] Zumberge J E. Prediction of source rock characteristics based on terpane biomarkers in crude oils: A multivariate statistical approach[J]. Geochimica et Cosmochimica Acta, 1987, 51(6): 1625-1637.
[48] Peters K E, Moldowan J M, Driscole A R, et al. Origin of Beatrice oil by co-sourcing from Devonian and Middle Jurassic source rocks, Inner Moray Firth, United Kingdom[J]. AAPG Bulletin, 1989, 73(4): 454-471.
[49] 敖添,王作栋,张婷,等. 塔河油田原油三环萜烷异常分布特征及成因探讨[J]. 沉积学报,2023,41(4):1202-1211.

Ao Tian, Wang Zuodong, Zhang Ting, et al. Abnormal distribution of tricyclic terpanes and its genesis in crude oils from Tahe oilfield[J]. Acta Sedimentologica Sinica, 2023, 41(4): 1202-1211.
[50] Erlich R N, Astorga A, Sofer Z, et al. Palaeoceanography of organic-rich rocks of the Loma Chumico Formation of Costa Rica, Late Cretaceous, eastern Pacific[J]. Sedimentology, 1996, 43(4): 691-718.
[51] Didyk B M, Simoneit B R T, Brassell S C, et al. Organic geochemical indicators of palaeoenvironmental conditions of sedimentation[J]. Nature, 1978, 272(5650): 216-222.
[52] Ma J, Huang Z L, Gao X Y, et al. Oil-source rock correlation for tight oil in tuffaceous reservoirs in the Permian Tiaohu Formation, Santanghu Basin, northwest China[J]. Canadian Journal of Earth Sciences, 2015, 52(11): 1014-1026.
[53] Ten H H L, Rohmer M, Rullkötter J, et al. Tetrahymanol, the most likely precursor of gammacerane, occurs ubiquitously in marine sediments[J]. Geochimica et Cosmochimica Acta, 1989, 53(11): 3073-3079.
[54] 陈治军,高怡文,刘护创,等. 银根—额济纳旗盆地哈日凹陷下白垩统烃源岩地球化学特征与油源对比[J]. 石油学报,2018,39(1):69-81.

Chen Zhijun, Gao Yiwen, Liu Huchuang, et al. Geochemical characteristics of Lower Cretaceous source rocks and oil-source correlation in Hari Sag, Yingen-Ejinaqi Basin[J]. Acta Petrolei Sinica, 2018, 39(1): 69-81.
[55] Li Y, Hou D J, Cheng X, et al. Geochemical characteristics and significance of aromatic hydrocarbons in crude oil from the east Fukang Sag, Junggar Basin, NW China[J]. Frontiers in Earth Science, 2023, 10: 1031367.
[56] Huang D F, Li J C, Zhang D J. Maturation sequence of continental crude oils in hydrocarbon basins in China and its significance[J]. Organic Geochemistry, 1990, 16(1/2/3): 521-529.
[57] Seifert W K, Moldowan J M. Use of biological markers in petroleum exploration[J]. Methods in Geochemistry and Geophysics, 1986, 24: 261-290.
[58] Peters K E, Moldowan J M. Effects of source, thermal maturity, and biodegradation on the distribution and isomerization of homohopanes in petroleum[J]. Organic Geochemistry, 1991, 17(1): 47-61.
[59] Seifert W K, Moldowan J M. Effect of biodegradation on steranes and terpanes in crude oils[J]. Geochimica et Cosmochimica Acta, 1979, 43(1): 111-126.
[60] Jiang Z S, Fowler M G. Carotenoid-derived alkanes in oils from northwestern China[J]. Organic Geochemistry, 1986, 10(4/5/6): 831-839.
[61] Wang Q R, Huang H P, Li Z, et al. Geochemical significance of β-carotane in lacustrine oils from the Shahejie Formation of the Dongying Depression, eastern China[J]. Organic Geochemistry, 2021, 156: 104241.
[62] Wang D Y, Li M J, Chen J F, et al. The biological source and paleoenvironment implication of rare short-chain lanostanes in alkaline lacustrine sediments[J]. Organic Geochemistry, 2023, 186: 104705.
[63] Boudou J P, Trichet J, Robinson N, et al. Profile of aliphatic hydrocarbons in a recent Polynesian microbial mat[J]. International Journal of Environmental Analytical Chemistry, 1986, 26(2): 137-155.
[64] Köster J, Volkman J K, Rullkötter J, et al. Mono-, di- and trimethyl-branched alkanes in cultures of the filamentous cyanobacterium Calothrix scopulorum[J]. Organic Geochemistry, 1999, 30(11): 1367-1379.
[65] Shiea J, Brassell S C, Ward D M. Mid-chain branched mono- and dimethyl alkanes in hot spring cyanobacterial mats: A direct biogenic source for branched alkanes in ancient sediments?[J]. Organic Geochemistry, 1990, 15(3): 223-231.
[66] Hou M G, Zha M, Liu H, et al. The coupling control of biological precursors and environmental factors on β-carotane enrichment in alkaline lacustrine source rocks: A case study from the Fengcheng Formation in the western Junggar Basin, NW China[J]. Petroleum Science, 2024, doi:  10.1016/j.petsci.2023.12.019.
[67] Xia L W, Cao J, Hu W X, et al. Coupling of paleoenvironment and biogeochemistry of deep-time alkaline lakes: A lipid biomarker perspective[J]. Earth-Science Reviews, 2021, 213: 103499.
[68] 支东明,李建忠,陈旋,等. 吐哈探区深层油气勘探进展及潜力评价[J]. 新疆石油地质,2023,44(3):253-264.

Zhi Dongming, Li Jianzhong, Chen Xuan, et al. Exploration progress and potential evaluation of deep oil and gas in Turpan-Hami exploration area[J]. Xinjiang Petroleum Geology, 2023, 44(3): 253-264.
[69] 支东明,唐勇,郑孟林,等. 准噶尔盆地玛湖凹陷风城组页岩油藏地质特征与成藏控制因素[J]. 中国石油勘探,2019,24(5):615-623.

Zhi Dongming, Tang Yong, Zheng Menglin, et al. Geological characteristics and accumulation controlling factors of shale reservoirs in Fengcheng Formation, Mahu Sag, Junggar Basin[J]. China Petroleum Exploration, 2019, 24(5): 615-623.
[70] 支东明,唐勇,何文军,等. 准噶尔盆地玛湖凹陷风城组常规—非常规油气有序共生与全油气系统成藏模式[J]. 石油勘探与开发,2021,48(1):38-51.

Zhi Dongming, Tang Yong, He Wenjun, et al. Orderly coexistence and accumulation models of conventional and unconventional hydrocarbons in Lower Permian Fengcheng Formation, Mahu Sag, Junggar Basin[J]. Petroleum Exploration and Development, 2021, 48(1): 38-51.
[71] Wang W H, Cui H S, Tan J Q, et al. Permian cyanobacterial blooms resulted in enrichment of organic matter in the Lucaogou Formation in the Junggar Basin, NW China[J]. Minerals, 2023, 13(4): 537.
[72] Zhang S H, Liu C Y, Liang H, et al. Mineralogical composition and organic matter characteristics of lacustrine fine-grained volcanic-hydrothermal sedimentary rocks: A data-driven analytics for the Second member of Permian Lucaogou Formation, Santanghu Basin, NW China[J]. Marine and Petroleum Geology, 2021, 126: 104920.
[73] Brocks J J, Buick R, Summons R E, et al. A reconstruction of Archean biological diversity based on molecular fossils from the 2.78 to 2.45 billion-year-old Mount Bruce Supergroup, Hamersley Basin, western Australia[J]. Geochimica et Cosmochimica Acta, 2003, 67(22): 4321-4335.
[74] 张俊杰,邢磊,侯笛. 蓝细菌生物标志物:2-甲基藿烷类化合物研究进展及其应用[J]. 地球环境学报,2018,9(5):434-443.

Zhang Junjie, Xing Lei, Hou Di. Progress of research on the cyanobacterial biomarker: 2-methylhopanoid and its application[J]. Journal of Earth Environment, 2018, 9(5): 434-443.
[75] Grba N, Šajnović A, Stojanović K, et al. Preservation of diagenetic products of β-carotene in sedimentary rocks from the Lopare Basin (Bosnia and Herzegovina)[J]. Geochemistry, 2014, 74(1): 107-123.
[76] Xie X M, Borjigin T, Zhang Q Z, et al. Intact microbial fossils in the Permian Lucaogou Formation oil shale, Junggar Basin, NW China[J]. International Journal of Coal Geology, 2015, 146: 166-178.
[77] Kuang L C, Tang Y, Lei D W, et al. Formation conditions and exploration potential of tight oil in the Permian saline lacustrine dolomitic rock, Junggar Basin, NW China[J]. Petroleum Exploration and Development, 2012, 39(6): 700-711.
[78] Liang P, Chen L, Li R C, et al. In-situ element geochemical and sulfur isotope signature of pyrite and chalcopyrite: Constraints on ore-forming processes of the Laoshankou iron oxide-copper (-gold) deposit, northern East Junggar[J]. Ore Geology Reviews, 2021, 139: 104510.